…The researchers’ reconstruction of this animal’s footfall sequence reveals a distinctive gait called a lateral-sequence walk, in which the legs on one side of the animal move in succession, the rear leg followed by the foreleg, alternating with the movement of the two legs on the opposite side. “Living species of tetrapods―dogs and cats, for example―routinely use a lateral-sequence gait when they walk slowly,” says Rowland. “The Bright Angel Trail tracks document the use of this gait very early in the history of vertebrate animals. We previously had no information about that.” Also revealed by the trackways is the earliest-known utilization of sand dunes by vertebrate animals. (Cliff Collapse Reveals 313-million-year-old Fossil Footprints in Grand Canyon National Park)